OpenGLException in TestToneMapFilter

While running TestToneMapFilter in jme3-examples, press the spacebar twice:

May 13, 2021 2:28:56 PM com.jme3.system.JmeDesktopSystem initialize
INFO: Running on jMonkeyEngine 3.4.0-SNAPSHOT
 * Branch: v3.4
 * Git Hash: bdf15ba
 * Build Date: 2021-05-13
May 13, 2021 2:28:57 PM com.jme3.system.lwjgl.LwjglContext printContextInitInfo
INFO: LWJGL 2.9.3 context running on thread jME3 Main
 * Graphics Adapter: null
 * Driver Version: null
 * Scaling Factor: 1
May 13, 2021 2:28:57 PM com.jme3.renderer.opengl.GLRenderer loadCapabilitiesCommon
INFO: OpenGL Renderer Information
 * Vendor: NVIDIA Corporation
 * Renderer: GeForce GT 545/PCIe/SSE2
 * OpenGL Version: 4.6.0 NVIDIA 390.143
 * GLSL Version: 4.60 NVIDIA
 * Profile: Compatibility
May 13, 2021 2:28:57 PM com.jme3.audio.openal.ALAudioRenderer initOpenAL
INFO: Audio Renderer Information
 * Device: OpenAL Soft
 * Vendor: OpenAL Community
 * Renderer: OpenAL Soft
 * Version: 1.1 ALSOFT 1.15.1
 * Supported channels: 64
 * ALC extensions: ALC_ENUMERATE_ALL_EXT ALC_ENUMERATION_EXT ALC_EXT_CAPTURE ALC_EXT_DEDICATED ALC_EXT_disconnect ALC_EXT_EFX ALC_EXT_thread_local_context ALC_SOFT_loopback
 * AL extensions: AL_EXT_ALAW AL_EXT_DOUBLE AL_EXT_EXPONENT_DISTANCE AL_EXT_FLOAT32 AL_EXT_IMA4 AL_EXT_LINEAR_DISTANCE AL_EXT_MCFORMATS AL_EXT_MULAW AL_EXT_MULAW_MCFORMATS AL_EXT_OFFSET AL_EXT_source_distance_model AL_LOKI_quadriphonic AL_SOFT_buffer_samples AL_SOFT_buffer_sub_data AL_SOFTX_deferred_updates AL_SOFT_direct_channels AL_SOFT_loop_points AL_SOFT_source_latency
May 13, 2021 2:28:57 PM com.jme3.audio.openal.ALAudioRenderer initOpenAL
WARNING: Pausing audio device not supported.
May 13, 2021 2:28:57 PM com.jme3.audio.openal.ALAudioRenderer initOpenAL
INFO: Audio effect extension version: 1.0
May 13, 2021 2:28:57 PM com.jme3.audio.openal.ALAudioRenderer initOpenAL
INFO: Audio max auxiliary sends: 4
== Tone Mapping Sample ==
 SPACE:	Toggle tone-mapping OFF or ON
 Y:	Increase white-point
 H:	Decrease white-point
May 13, 2021 2:28:58 PM com.jme3.material.Material checkTextureParamColorSpace
WARNING: The texture Textures/HdrTest/Memorial.hdr has linear color space, but the material parameter ColorMap specifies no color space requirement, this may lead to unexpected behavior.
Check if the image was not set to another material parameter with a linear color space, or that you did not set the ColorSpace to Linear using texture.getImage.setColorSpace().
Tone Mapping OFF
Tone Mapping ON
May 13, 2021 2:29:00 PM com.jme3.app.LegacyApplication handleError
SEVERE: An OpenGL error has occurred!
org.lwjgl.opengl.OpenGLException: Invalid operation (1282)
	at org.lwjgl.opengl.Util.checkGLError(Util.java:59)
	at com.jme3.system.lwjgl.LwjglAbstractDisplay.checkGLError(LwjglAbstractDisplay.java:145)
	at com.jme3.system.lwjgl.LwjglAbstractDisplay.runLoop(LwjglAbstractDisplay.java:166)
	at com.jme3.system.lwjgl.LwjglDisplay.runLoop(LwjglDisplay.java:201)
	at com.jme3.system.lwjgl.LwjglAbstractDisplay.run(LwjglAbstractDisplay.java:242)
	at java.lang.Thread.run(Thread.java:748)

After enabling graphics debug:

Tone Mapping OFF
Tone Mapping ON
[JME3] OpenGL debug message
       ID: 1282
       Source: API
       Type: ERROR
       Severity: HIGH
       Message: GL_INVALID_OPERATION error generated. Render buffer not bound.
java.lang.Exception: Stack trace
	at java.lang.Thread.dumpStack(Thread.java:1336)
	at com.jme3.system.lwjgl.LwjglGLDebugOutputHandler.handleMessage(LwjglGLDebugOutputHandler.java:76)
	at org.lwjgl.opengl.GL30.nglRenderbufferStorage(Native Method)
	at org.lwjgl.opengl.GL30.glRenderbufferStorage(GL30.java:1079)
	at com.jme3.renderer.lwjgl.LwjglGLFboGL3.glRenderbufferStorageEXT(LwjglGLFboGL3.java:94)
	at sun.reflect.NativeMethodAccessorImpl.invoke0(Native Method)
	at sun.reflect.NativeMethodAccessorImpl.invoke(NativeMethodAccessorImpl.java:62)
	at sun.reflect.DelegatingMethodAccessorImpl.invoke(DelegatingMethodAccessorImpl.java:43)
	at java.lang.reflect.Method.invoke(Method.java:498)
	at com.jme3.renderer.opengl.GLDebug.invoke(GLDebug.java:107)
	at com.sun.proxy.$Proxy2.glRenderbufferStorageEXT(Unknown Source)
	at com.jme3.renderer.opengl.GLRenderer.updateRenderBuffer(GLRenderer.java:1812)
	at com.jme3.renderer.opengl.GLRenderer.updateFrameBufferAttachment(GLRenderer.java:1869)
	at com.jme3.renderer.opengl.GLRenderer.updateFrameBuffer(GLRenderer.java:1922)
	at com.jme3.renderer.opengl.GLRenderer.setFrameBuffer(GLRenderer.java:2076)
	at com.jme3.renderer.RenderManager.renderViewPort(RenderManager.java:1102)
	at com.jme3.renderer.RenderManager.render(RenderManager.java:1180)
	at com.jme3.app.SimpleApplication.update(SimpleApplication.java:273)
	at com.jme3.system.lwjgl.LwjglAbstractDisplay.runLoop(LwjglAbstractDisplay.java:160)
	at com.jme3.system.lwjgl.LwjglDisplay.runLoop(LwjglDisplay.java:201)
	at com.jme3.system.lwjgl.LwjglAbstractDisplay.run(LwjglAbstractDisplay.java:242)
	at java.lang.Thread.run(Thread.java:748)
May 13, 2021 2:30:58 PM com.jme3.app.LegacyApplication handleError
SEVERE: Uncaught exception thrown in Thread[jME3 Main,5,main]
com.jme3.renderer.RendererException: An OpenGL error occurred - Invalid operation (Error Code: 1282)
	at com.jme3.renderer.opengl.GLDebug.checkError(GLDebug.java:101)
	at com.jme3.renderer.opengl.GLDebug.invoke(GLDebug.java:113)
	at com.sun.proxy.$Proxy2.glRenderbufferStorageEXT(Unknown Source)
	at com.jme3.renderer.opengl.GLRenderer.updateRenderBuffer(GLRenderer.java:1812)
	at com.jme3.renderer.opengl.GLRenderer.updateFrameBufferAttachment(GLRenderer.java:1869)
	at com.jme3.renderer.opengl.GLRenderer.updateFrameBuffer(GLRenderer.java:1922)
	at com.jme3.renderer.opengl.GLRenderer.setFrameBuffer(GLRenderer.java:2076)
	at com.jme3.renderer.RenderManager.renderViewPort(RenderManager.java:1102)
	at com.jme3.renderer.RenderManager.render(RenderManager.java:1180)
	at com.jme3.app.SimpleApplication.update(SimpleApplication.java:273)
	at com.jme3.system.lwjgl.LwjglAbstractDisplay.runLoop(LwjglAbstractDisplay.java:160)
	at com.jme3.system.lwjgl.LwjglDisplay.runLoop(LwjglDisplay.java:201)
	at com.jme3.system.lwjgl.LwjglAbstractDisplay.run(LwjglAbstractDisplay.java:242)
	at java.lang.Thread.run(Thread.java:748)